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Salmon
is the common name for several species of fish
of the family Salmonidae.
Several other fish in the family are called trout.
Salmon live in both the Atlantic
and Pacific
Oceans, as well as the Great
Lakes and other land locked lakes.
Typically,
salmon are anadromous:
they are born in fresh
water, migrate to the ocean, then return to fresh water to reproduce. Folklore
has it that the fish return to the exact spot where they were born to
spawn, and modern research shows that usually at least 90% of the fish
that spawn
in a particular stream were born there.
The precise method salmon use to navigate has not been entirely
established, though their keen sense of smell is certainly involved. In
all species of Pacific salmon, the mature individuals die within a few
days or weeks of spawning, a trait known as semelparity.
However, even in those species of salmon that may survive to spawn more
than once (iteroparity),
post-spawning mortality is quite high (perhaps as high as 40 to 50%.)
Those species average about two or, perhaps, three spawning events per
individual.
In
order to lay her roe,
the female salmon uses her tail fin to excavate a shallow depression,
called a redd. The redd may sometimes contain 5,000 eggs covering
30 square feet. The eggs usually range from orange to red in color. One or more males will
approach the female in her redd, depositing his sperm, or milt, over the
roe. The female then covers the eggs by disturbing the gravel at the upstream
edge of the depression before moving on to make another redd. The female
will make as many as 7 redds before her supply of eggs is exhausted. The
salmon then die within a few days of spawning.
The
eggs will hatch into alevin or sac fry. The fry quickly
develop into parr with camouflaging vertical stripes. The parr stay
for one to three years in their natal stream before becoming smolts
which are distinguished by their bright silvery colour with scales that
are easily rubbed off. It is estimated that only 10% of all salmon eggs
survive long enough to reach this stage. The smolt body chemistry changes, allowing them to live in salt water.
Smolts spend a portion of their out-migration time in brackish water,
where their body chemistry becomes accustomed to osmoregulation
in the ocean.
The
salmon spend one to five years (depending on the species) in the open
ocean where they will become sexually mature. The adult salmon returns
primarily to its natal stream to spawn. When fish return for the first
time they are called whitling in Scotland.
Prior to spawning, depending
on the species,
the salmon undergoes changes. They may grow a hump, develop canine teeth,
develop a kype (a pronounced curvature of the jaws in male salmon).
All will change from the silvery blue of a fresh run fish from the sea to
a darker color. Condition tends to deteriorate the longer the fish remain
in freshwater, and they then deteriorate further after they spawn becoming
known as kelts. Salmon can make amazing journeys, sometimes moving
hundreds of miles upstream against strong currents and rapids to
reproduce.
The
age of a salmon can be deduced from the growth rings on its scales,
examined under the microscope.
Each year, the fish experiences a period of rapid
growth, often in summer, and one of slower growth, normally in winter.
This results in rings (annuli) analogous to the growth rings visible in a
tree trunk. Freshwater growth shows as densely crowded rings, sea growth
as widely spaced rings; spawning is marked by significant erosion as body
mass is converted into eggs and milt.
Freshwater
streams and estuaries provide important habitat for many salmon species.
They feed on terrestrial
and aquatic
insects, amphipods,
and other crustaceans
while young, and primarily on other fish when older. Eggs are laid in
deeper water with larger gravel, and need cool water and good water flow
(to supply oxygen) to the developing embryos. Mortality of salmon in the
early life stages is usually high due to natural predation and human
induced changes in habitat, such as siltation, high water temperatures,
low oxygen conditions, loss of stream cover, and reductions in river flow.
Estuaries
and their associated wetlands
provide vital nursery areas for the salmon prior to their departure to the
open ocean. Wetlands not only help buffer the estuary from silt and
pollutants, but also provide important feeding and hiding areas.
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